तदा then, सिंहव्याघ्रमृगद्विजाः lions, tigers, deer and birds, समन्तात् अभिसम्पत्य coming together from all sides, रोषात् due to anguish, छायानुगामिनः followers of the shadow, सीताम् Sita, अन्वधावन् ran following her.

Then the lions, tigers, deer and birds gathered together in anger (also anguish) and ran after Sita's shadow.