ते वानराः set his foot those monkeys, समुद्रेषु on the sea shore, गिरिषु on the mountains, वनेषु च in the forests, सरस्सु च on the banks of lakes, सर्वान् all over, वानरान् monkeys, रामहेतोः to serve Rama's cause, अचोदयन् driven.

The messenger monkeys drove in all those living on the sea shore, mountains, forests as well as banks of lakes in order to serve the cause of Rama.