अथ: thus, मुख्यज्ञः who know the important ones, सारणः Saarana, परिपृच्छतः being asked, राक्षसेन्द्रस्य: by the king of Rakshasas, मुख्यांस्तांस्तु: important there, वनौकसः of the wanderers in woods, वचनम् these words, आचचक्षे: speak out.

Hearing the words asked by Rakshasendra, Saarana, who knew the important ones, thus spoke out about the important monkey leaders.