तदा then, युयुत्सवः who were eager to combat, सर्वे all,ते of them, वानरगणाः vanara groups, समुत्पत्य jumped down, गुहाभ्यः from the caves, शिखरेभ्यश्च from mountain peaks, आशु from trees, पुप्लुविरे heroic monkeys

Then all the heroic monkeys who were eager for combat jumped down from the caves, mountain peaks and trees.